Studies nonhuman primates. Researches human evolution. Specializes in mammalian taxonomy. Focuses on biogeography. Examines animal domestication. Conducted museum work on primates worldwide. Lifelong studies on primate classification variation evolution. Published Primate Taxonomy book 2001. Continues research with new discoveries. Invited to primatologist conferences. Spoke at Goettingen Freilandtage 2005. Attended African Genesis symposium 2006. Publishes on Homo floresiensis. Paper in press with PhD student Debbie Argue. Studies ungulate taxonomy. Researches elephants carnivores. Biogeography reconstructs past climates geography. Fieldwork in Kenya Tanzania Rwanda India Iran China Indonesia. Recent fieldwork in Sri Lanka Democratic Republic Congo. Collaborates with Europe China Sri Lanka colleagues. Much research sole-authored.