Basal Eurasian ancestry reached 66% in Mesolithic Iran and 44% in Natufian Levant. Basal Eurasians split from other non-Africans before Ust'-Ishim and Han Chinese divergence. Basal Eurasian ancestry correlates negatively with Neanderthal admixture, implying Basal Eurasians had none. Levantine farmers descended from local Natufian hunter-gatherers. Iranian farmers descended from local Iranian and Caucasus hunter-gatherers. Early Near Eastern farmers showed extreme genetic differentiation like between West and East Eurasians. Bronze Age admixture homogenized Near East populations to modern low Fst levels. Anatolian-related farmers spread ancestry to Europe. Levantine-related farmers spread ancestry to East Africa around 1000 BCE. Iran Chalcolithic-related ancestry contributed 43% to Eurasian steppe Bronze Age. Iran and steppe ancestries mixed into South Asian Ancestral North Indians. Natufians carried African-origin Y-haplogroup E but no sub-Saharan autosomal affinity. West Eurasians form mixtures of four ancient ancestry streams: WHG, EHG, Levant Natufian, Iran Neolithic. East Asians carry 5-10% Ancient North Eurasian ancestry. European farmers unrelated to Levantine farmers. Near East farming spread via local continuity, not population replacement.
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