986 skulls from Rwanda, 41 from Tanzania, 4 from Burundi, 54 from East Africa. Collected by anthropologist Felix von Luschan to study mankind development. Shipped from German East African colonies around 1907-08. Stored in deep storage through two world wars. Transferred to Prussian Cultural Heritage Foundation in 2011 from Charite hospital. Skulls in very poor condition with most paperwork lost. Notes on bones provide main clues to origins. Two-year study to determine provenance and possible return to Africa. Collaboration planned with Rwanda, Tanzania, Burundi researchers. Rwanda unaware of skulls until 2016. Rwandan ambassador questions how 1000 skulls obtained given no cemeteries then. Bodies traditionally buried near family homes in Rwanda. Germany previously returned 20 Namibian skulls from colonial genocide. Namibian skulls among 300 taken after anti-colonial uprising slaughter. Skulls gathered dust in German archives until 2008 exposure.