95 endogenous proteins retrieved from Harbin cranium dating to at least 146,000 years ago. Harbin individual has three Denisovan-derived amino acid variants. Harbin clusters with Denisova 3 confirming Denisovan population membership. Previously called Homo longi but now identified as Denisovan via proteins. Ancient proteomics identifies Denisovans where DNA fails. Bridges morphological and molecular evidence for Denisovans. Enables study of Denisovan dispersal and evolution. Harbin cranium from China shows Denisovans in East Asia long ago. Proteins degrade slower than DNA allowing older remains analysis. Denisovan fossils fragmentary but proteomics fills identification gap.
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